Peer-Reviewed Publications and Patents

John A. Byers

John Byers 2012



M.S., Department of Entomology, Colorado State University, Fort Collins, Colorado:

1. Byers, J.A., Brewer, J.W., and Denna, D.W. 1976. Plant growth hormones in pinyon insect galls. Marcellia 39:125-134. (Institut de botanique de Strasbourg)
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Ph.D. and Post-Doc, Department of Entomological Sciences, University of California, Berkeley, California:

2. Byers, J.A., Wood, D.L., Browne, L.E., Fish, R.H., Piatek, B., and Hendry, L.B. 1979. Relationship between a host plant compound, myrcene and pheromone production in the bark beetle, Ips paraconfusus. Journal of Insect Physiology 25(6):477-482.
DOI: https://doi.org/10.1016/S0022-1910(79)80005-0 pdf

3. Byers, J.A. and Wood, D.L. 1980. Interspecific inhibition of the response of the bark beetles, Dendroctonus brevicomis and Ips paraconfusus, to their pheromones in the field. Journal of Chemical Ecology 6(1):149-164.
DOI: https://doi.org/10.1007/BF00987534 pdf

4. Byers, J.A., Svihra, P., and Koehler, C.S. 1980. Attraction of elm bark beetles to cut limbs on elm. Journal of Arboriculture 6:245-246.
DOI: https://joa.isa-arbor.com/ pdf

5. Hendry, L.B., Piatek, B., Browne, L.E., Wood, D.L., Byers, J.A., Fish, R.H., and Hicks, R.A. 1980. In vivo conversion of a labelled host plant chemical to pheromones of the bark beetle Ips paraconfusus. Nature 284(5755):485.
DOI: https://doi.org/10.1038/284485a0 pdf

6. Byers, J.A. 1981a. Pheromone biosynthesis in the bark beetle, Ips paraconfusus, during feeding or exposure to vapours of host plant precursors. Insect Biochemistry 11(5):563-569.
DOI: https://doi.org/10.1016/0020-1790(81)90024-X pdf

7. Byers, J.A. 1981b. Effect of mating on terminating aggregation during host colonization in the bark beetle, Ips paraconfusus. Journal of Chemical Ecology 7(6):1135-1147.
DOI: https://doi.org/10.1007/BF00987633 pdf

8. Byers, J.A. 1981c. Versatile electronic timer for synchronous switching of multiple electrical devices. Behavior Research Methods and Instrumentation 13(3):381-383.
DOI: https://doi.org/10.3758/BF03202039 pdf

9. Byers, J.A. and Wood, D.L. 1981a. Interspecific effects of pheromones on the attraction of the bark beetles, Dendroctonus brevicomis and Ips paraconfusus in the laboratory. Journal of Chemical Ecology 7(1):9-18.
DOI: https://doi.org/10.1007/BF00988631 pdf

10. Byers, J.A. and Wood, D.L. 1981b. Antibiotic-induced inhibition of pheromone synthesis in a bark beetle. Science 213(4509):763-764.
DOI: https://doi.org/10.1126/science.213.4509.763 pdf

11. Byers, J.A. 1982. Male-specific conversion of the host plant compound, myrcene, to the pheromone, (+)-ipsdienol, in the bark beetle, Dendroctonus brevicomis. Journal of Chemical Ecology 8(2):363-371.
DOI: https://doi.org/10.1007/BF00987784 pdf

12. Byers, J.A., and Poinar, G.O., Jr. 1982. Location of insect hosts by the nematode, Neoaplectana carpocapsae, in response to temperature. Behaviour 79(1):1-10.
DOI: https://doi.org/10.1163/156853982X00535 pdf

13. Byers, J.A. 1983a. Influence of sex, maturity and host substances on pheromones in the guts of the bark beetles, Ips paraconfusus and Dendroctonus brevicomis. Journal of Insect Physiology 29(1):5-13.
DOI: https://doi.org/10.1016/0022-1910(83)90100-2 pdf

14. Byers, J.A. 1983b. Bark beetle conversion of a plant compound to a sex-specific inhibitor of pheromone attraction. Science 220(4597):624-626.
DOI: https://doi.org/10.1126/science.220.4597.624 pdf

15. Byers, J.A. 1983c. Sex-specific responses to aggregation pheromone: Regulation of colonization density in the bark beetle Ips paraconfusus. Journal of Chemical Ecology 9(1):129-142.
DOI: https://doi.org/10.1007/BF00987777 pdf

16. Byers, J.A. 1983d. Electronic fraction collector used for insect sampling in the photoperiod-induced diel emergence of bark beetles. Physiological Entomology 8(2):133-138.
DOI: https://doi.org/10.1007/BF01022541 pdf

17. Byers, J.A., and Unkrich, M.A. 1983. Electronic light intensity control to simulate dusk and dawn conditions. Annals of the Entomological Society of America 76(3):556-558.
DOI: https://doi.org/10.1093/aesa/76.3.556 pdf

Post-Doctoral, Chemical Ecology Group, Department of Animal Ecology, Lund University, Sweden:

18. Byers, J.A. 1984a. Electronic multiprobe thermometer and multiplexer for recording temperatures of microenvironments in the forest litter habitat of bark beetles (Coleoptera: Scolytidae). Environmental Entomology 13(3)863-867.
DOI: https://doi.org/10.1093/ee/13.3.863 pdf

19. Byers, J.A. 1984b. Nearest neighbor analysis and simulation of distribution patterns indicates an attack spacing mechanism in the bark beetle, Ips typographus (Coleoptera: Scolytidae). Environmental Entomology 13(5):1191-1200.
DOI: https://doi.org/10.1093/ee/13.5.1191 pdf

20. Byers, J.A., Wood, D.L., Craig, J. and Hendry, L.B. 1984. Attractive and inhibitory pheromones produced in the bark beetle, Dendroctonus brevicomis, during host colonization: Regulation of inter- and intraspecific competition. Journal of Chemical Ecology 10(6):861-877.
DOI: https://doi.org/10.1007/BF00987969 pdf

21. Byers, J.A., Lanne, B.S., Löfqvist, J., Schlyter, F. and Bergström, G. 1985. Olfactory recognition of host-tree susceptibility by pine shoot beetles. Naturwissenschaften 72(6):324-326.
DOI: https://doi.org/10.1007/BF00454776 pdf

22. Byers, J.A. 1987a. Interactions of pheromone component odor plumes of western pine beetle. Journal of Chemical Ecology 13(12):2143-2157.
DOI: https://doi.org/10.1007/BF01012563 pdf

23. Byers, J.A. 1987b. Novel fraction collector for studying the oviposition rhythm in the turnip moth. Chronobiology International 4(2):189-194.
DOI: https://doi.org/10.3109/07420528709078525 pdf

24. Byers, J., Birgersson, G., Löfqvist, J. and Bergström. 1987. A method and composition for observation and control of Pityogenes chalcographus. European Patent #87850032.1.

25. Lanne, B.S., Schlyter, F., Byers, J.A., Löfqvist, J., Leufvén, A., Bergström, G., Van Der Pers, J.N.C., Unelius, R., Baeckström, P. and Norin, T. 1987. Differences in attraction to semiochemicals present in sympatric pine shoot beetles, Tomicus minor and T. piniperda. Journal of Chemical Ecology 13(5):1045-1067.
DOI: https://doi.org/10.1007/BF01020537 pdf

26. Schlyter, F., Byers, J.A. and Löfqvist, J. 1987a. Attraction to pheromone sources of different quantity, quality, and spacing: Density-regulation mechanisms in bark beetle Ips typographus. Journal of Chemical Ecology 13(6):1503-1523.
DOI: https://doi.org/10.1007/BF01012294 pdf

27. Schlyter, F., Löfqvist, J. and Byers, J.A. 1987b. Behavioural sequence in attraction of the bark beetle Ips typographus to pheromone sources. Physiological Entomology 12(2):185-196.
DOI: https://doi.org/10.1111/j.1365-3032.1987.tb00741.x pdf

28. Schlyter, F., Birgersson, G., Byers, J.A., Löfqvist, J. and Bergström, G. 1987c. Field response of the spruce bark beetle, Ips typographus, to aggregation pheromone candidates. Journal of Chemical Ecology 13(4):701-716.
DOI: https://doi.org/10.1007/BF01020153 pdf

29. Byers, J.A. 1988a. Upwind flight orientation to pheromone in western pine beetle tested with rotating windvane traps. Journal of Chemical Ecology 14(1):189-198.
DOI: https://doi.org/10.1007/BF01022541 pdf

30. Byers, J.A. 1988b. Novel diffusion-dilution method for release of semiochemicals: Testing pheromone component ratios on western pine beetle. Journal of Chemical Ecology 14(1):199-212.
DOI: https://doi.org/10.1007/BF01022542 pdf

31. Byers, J.A., Birgersson, G., Löfqvist, J., and Bergström, G. 1988. Synergistic pheromones and monoterpenes enable aggregation and host recognition by a bark beetle, Pityogenes chalcographus. Naturwissenschaften 75(3):153-155.
DOI: https://doi.org/10.1007/BF00405312 pdf

32. Byers, J.A. 1989a. Chemical ecology of bark beetles. Experientia 45(3):271-283.
DOI: https://doi.org/10.1007/BF01951813 pdf

33. Byers, J.A. 1989b. Behavioral mechanisms involved in reducing competition in bark beetles. Ecography 12(4):466-476.
DOI: https://doi.org/10.1111/j.1600-0587.1989.tb00924.x pdf

34. Byers, J. and Löfqvist, J. 1989. A method and composition for protection of forest products from spruce bark beetle, Ips typographus. Swedish Patent W08911221.

35. Byers, J.A. and Löfqvist, J. 1989. Flight initiation and survival in the bark beetle Ips typographus (Coleoptera: Scolytidae) during the spring dispersal. Ecography 12(4):432-440.
DOI: https://doi.org/10.1111/j.1600-0587.1989.tb00919.x pdf

36. Byers, J.A., Anderbrant, O., and Löfqvist, J. 1989a. Effective attraction radius: A method for comparing species attractants and determining densities of flying insects. Journal of Chemical Ecology 15(2):749-765.
DOI: https://doi.org/10.1007/BF01014716 pdf

37. Byers, J.A., Högberg, H.E., Unelius, R., Birgersson, G. and Löfqvist, J. 1989b. Structure-activity studies on aggregation pheromone components of Pityogenes chalcographus (Coleoptera: Scolytidae): all stereoisomers of chalcogran and methyl 2,4-decadienoate. Journal of Chemical Ecology 15(2):685-695.
DOI: https://doi.org/10.1007/BF01014711 pdf

38. Byers, J.A., Lanne, B.S., and Löfqvist, J. 1989c. Host-tree unsuitability recognized by pine shoot beetles in flight. Experientia 45(5):489-492.
DOI: https://doi.org/10.1007/BF01952042 pdf

39. Byers, J., Lanne, B., Löfqvist, J. and Schlyter, F. 1989. A method and composition for protection of forest products from pine shoot beetles, Tomicus piniperda and T. minor. Swedish Patent W08910692.

Assistant Professor, Department of Animal Ecology, Lund University, Sweden:

40. Birgersson, G., Byers, J.A., Bergström, G., and Löfqvist, J. 1990. Production of pheromone components, chalcogran and methyl (E,Z)-2,4- decadienoate, in the spruce engraver Pityogenes chalcographus. Journal of Insect Physiology 36(6):391-395.
DOI: https://doi.org/10.1016/0022-1910(90)90056-L pdf

41. Byers, J.A., and Birgersson, G. 1990. Pheromone production in a bark beetle independent of myrcene precursor in host pine species. Naturwissenschaften 77(8):385-387.
DOI: https://doi.org/10.1007/BF01135739 pdf

42. Byers, J.A., Birgersson, G., Löfqvist, J., Appelgren, M., and Bergström, G. 1990a. Isolation of pheromone synergists of bark beetle, Pityogenes chalcographus, from complex insect-plant odors by fractionation and subtractive-combination bioassay. Journal of Chemical Ecology 16(3):861-876.
DOI: https://doi.org/10.1007/BF01016496 pdf

43. Byers, J.A., Schlyter, F., Birgersson, G., and Francke, W. 1990b. E- Myrcenol in Ips duplicatus: An aggregation pheromone component new for bark beetles. Experientia 46(11):1209-1211.
DOI: https://doi.org/10.1007/BF01936939 pdf

44. Byers, J.A. 1991a. Simulation of mate-finding behaviour of pine shoot beetles, Tomicus piniperda. Animal Behaviour 41(4):649-660.
DOI: https://doi.org/10.1016/S0003-3472(05)80903-0 pdf

45. Byers, J.A. 1991b. Pheromones and chemical ecology of locusts. Biological Reviews 66(4):347-378.
DOI: https://doi.org/10.1111/j.1469-185X.1991.tb01146.x pdf

46. Byers, J.A. 1991c. BASIC algorithms for random sampling and treatment randomization. Computers in Biology and Medicine 21(1):69-77.
DOI: https://doi.org/10.1016/0010-4825(91)90037-A pdf

47. Byers, J.A. 1991d. Cardcat: A scientific reference manager for IBM- compatible personal computers. American Entomologist 37(4):240-242.
DOI: https://doi.org/10.1093/ae/37.4.240 pdf

48. Byers, J.A. 1992a. Grid cell contour mapping of point densities: bark beetle attacks, fallen pine shoots, and infested trees. Oikos 63(2):233-243.
DOI: https://doi.org/10.2307/3545383 pdf

49. Byers, J.A. 1992b. Dirichlet tessellation of bark beetle spatial attack points. Journal of Animal Ecology 61(3):759-768.
DOI: https://doi.org/10.2307/5629 pdf

50. Byers, J.A. 1992c. Optimal fractionation and bioassay plans for isolation of synergistic chemicals: the subtractive-combination method. Journal of Chemical Ecology 18(9):1603-1621.
DOI: https://doi.org/10.1007/BF00993233 pdf

51. Byers, J.A. 1992d. Attraction of bark beetles, Tomicus piniperda, Hylurgops palliatus, and Trypodendron domesticum and other insects to short-chain alcohols and monoterpenes. Journal of Chemical Ecology 18(12):2385-2402.
DOI: https://doi.org/10.1007/BF00984957 pdf

52. Byers, J.A. 1992e. Trends in chemical ecology revealed with a personal computer program for searching data bases of scientific references and abstracts. Journal of Chemical Ecology 18(9):1481-1495.
DOI: https://doi.org/10.1007/BF00993222 pdf

53. Byers, J.A. 1992f. Ref-list: Program to list references found in DOS text files of scientific manuscripts. Bioinformatics 8(2):196-198.
DOI: https://doi.org/10.1093/bioinformatics/8.2.196 pdf

54. Schlyter, F., Birgersson, G., Byers, J.A., and Bakke, A. 1992. The aggregation pheromone of Ips duplicatus and its role in competitive interactions with I. typographus (Coleoptera: Scolytidae). Chemoecology 3(3):103-112.
DOI: https://doi.org/10.1007/BF01370137 pdf

55. Zhang, Q.H., Byers, J.A., and Schlyter, F. 1992. Optimal attack density in the larch bark beetle, Ips cembrae (Coleoptera: Scolytidae). Journal of Applied Ecology 29(3):672-678.
DOI: https://doi.org/10.2307/2404475 pdf

56. Byers, J.A. 1993a. Avoidance of competition by spruce bark beetles, Ips typographus and Pityogenes chalcographus. Experientia 49(3):272-275.
DOI: https://doi.org/10.1007/BF01923539 pdf

57. Byers, J.A. 1993b. Simulation and equation models of insect population control by pheromone-baited traps. Journal of Chemical Ecology 19(9):1939-1956.
DOI: https://doi.org/10.1007/BF00983798 pdf

58. Byers, J.A. 1993c. Orientation of bark beetles Pityogenes chalcographus and Ips typographus to pheromone-baited puddle traps placed in grids: A new trap for control of scolytids. Journal of Chemical Ecology 19(10):2297-2316.
DOI: https://doi.org/10.1007/BF00979665 pdf

59. Byers, J.A. 1993d. Randomization algorithms in BASIC for experimental design. Computers in Biology and Medicine 23(2):167-176.
DOI: https://doi.org/10.1016/0010-4825(93)90148-T pdf

60. Byers, J.A. 1993e. Conversion of BIOSIS Previews database files to reference text files and reprint request forms. American Entomologist 39(4):245-248.
DOI: https://doi.org/10.1093/ae/39.4.245 pdf

61. Byers, J. and Löfqvist, J. 1993. A method and composition for protection of forest products from spruce bark beetle, Pityogenes chalcographus. Swedish Patent US5223542.

62. Zhang, Q.H., Byers, J.A., and Zhang, X.D. 1993. Influence of bark thickness, trunk diameter and height on reproduction of the long- horned beetle, Monochamus sutor (Col., Cerambycidae) in burned larch and pine. Journal of Applied Entomology 115(1-5):145-154.
DOI: https://doi.org/10.1111/j.1439-0418.1993.tb00374.x pdf

63. Zolubas, P. and Byers, J.A., 1995. Recapture of dispersing bark beetles Ips typographus L. (Col., Scolytidae) in pheromone-baited traps. Journal of Applied Entomology 119(1-5):285-289.
DOI: https://doi.org/10.1111/j.1439-0418.1995.tb01287.x pdf

Researcher, Department of Plant Protection, Swedish University of Agricultural Science, Alnarp:

64. Byers, J.A. 1996a. An encounter rate model of bark beetle populations searching at random for susceptible host trees. Ecological Modelling 91(1):57-66.
DOI: https://doi.org/10.1016/0304-3800(95)00166-2 pdf

65. Byers, J.A. 1996b. Correct calculation of Dirichlet polygon areas. Journal of Animal Ecology 65(4):528-529.
DOI: https://doi.org/10.2307/5787 pdf

66. Byers, J.A. 1996c. Temporal clumping of bark beetle arrival at pheromone traps: Modeling anemotaxis in chaotic plumes. Journal of Chemical Ecology 22(11):2133-2155.
DOI: https://doi.org/10.1007/BF02040099 pdf

67. Byers, J.A. 1996d. Random selection algorithms for spatial and temporal sampling. Computers in Biology and Medicine 26(1):41-52.
DOI: https://doi.org/10.1016/0010-4825(95)00045-3 pdf

68. Byers, J.A. 1996e. Taxokey: a universal taxonomic key program using DOS text files and graphics. Bioinformatics 12(3):185-189.
DOI: https://doi.org/10.1093/bioinformatics/12.3.185 pdf

69. Byers, J.A., Zhang, Q.H., Schlyter, F., and Birgersson, G. 1998. Volatiles from nonhost birch trees inhibit pheromone response in spruce bark beetles. Naturwissenschaften 85(11):557-561.
DOI: https://doi.org/10.1007/s001140050551 pdf

70. Byers, J.A. 1999a. Effects of attraction radius and flight paths on catch of scolytid beetles dispersing outward through rings of pheromone traps. Journal of Chemical Ecology 25(5):985-1005.
DOI: https://doi.org/10.1023/A:1020869422943 pdf

71. Byers, J.A. 1999b. Database program to manage slides and images for teaching and presentations. Educational Media International 36(1):77-80.
DOI: https://doi.org/10.1080/0952398990360114 pdf

72. Byers, J.A. 1999c. Flash cards and animation software for education. Educational Media International 36(2):164-167.
DOI: https://doi.org/10.1080/0952398990360213 pdf

73. Byers, J.A. 1999d. Interactive learning using expert system quizzes on the internet. Educational Media International 36(3):191-194.
DOI: https://doi.org/10.1080/0952398990360304 pdf

74. Byers, J.A. 2000. Wind-aided dispersal of simulated bark beetles flying through forests. Ecological Modelling 125(2):231-243.
DOI: https://doi.org/10.1016/S0304-3800(99)00187-8 pdf

75. Byers, J.A., Zhang, Q.H., and Birgersson, G. 2000. Strategies of a bark beetle, Pityogenes bidentatus, in an olfactory landscape. Naturwissenschaften 87(11):503-507.
DOI: https://doi.org/10.1007/s001140050768 pdf

76. El-Sayed, A.M. and Byers, J.A. 2000. Inhibitory effect of monoterpenes on response of Pityogenes bidentatus to aggregation pheromone released by piezoelectric sprayer for precision release of semiochemicals. Journal of Chemical Ecology 26(8):1795-1809.
DOI: https://doi.org/10.1023/A:1005536421703 pdf

77. Schlyter, F., Zhang, Q-H., Anderson, P., Byers, J.A., Wadhams, L.J., Löfqvist, J., and Birgersson, G. 2000. Electrophysiological and behavioural responses of Scandinavian Tomicus piniperda and T. minor (Coleoptera: Scolytidae), to non-host leaf and bark volatiles. Canadian Entomologist 132(6):965-981.
DOI: https://doi.org/10.4039/Ent132965-6 pdf

78. Byers, J.A. 2001. Correlated random walk equations of animal dispersal resolved by simulation. Ecology 82(6):1680-1690.
DOI: https://doi.org/10.1890/0012-9658(2001)082%5B1680:CRWEOA%5D2.0.CO;2 pdf

Research Entomologist, Pest Management and Biological Control Research Unit, USDA-ARS, Phoenix / Maricopa, Arizona:

79. Byers, J.A. 2002. Internet programs for drawing moth pheromone analogs and searching literature database. Journal of Chemical Ecology 28(4):807-817.
DOI: https://doi.org/10.1023/A:1015244911584 pdf

80. Byers, J.A. 2004. Equations for nickel-chromium wire heaters of column transfer lines in gas chromatographic-electroantennographic detection (GC-EAD). Journal of Neuroscience Methods 135(1):89-93.
DOI: https://doi.org/10.1016/j.jneumeth.2003.12.004 pdf

81. Byers, J.A., Zhang, Q.H., and Birgersson, G. 2004. Avoidance of nonhost plants by a bark beetle, Pityogenes bidentatus, in a forest of odors. Naturwissenschaften 91(5):215-219.
DOI: https://doi.org/10.1007/s00114-004-0520-1 pdf

82. Blackmer, J.L., Rodriquez-Saona, C., Byers, J. A., Shope, K.L., and Smith, J.P. 2004. Behavioral response of Lygus hesperus to conspecifics and headspace volatiles of alfalfa in a Y-tube olfactometer. Journal of Chemical Ecology 30(8):1547-1564.
DOI: https://doi.org/10.1023/B:JOEC.0000042067.27698.30 pdf

83. Byers, J.A. 2005a. A cost of alarm pheromone production in cotton aphids, Aphis gossypii. Naturwissenschaften 92(2):69-72.
DOI: https://doi.org/10.1007/s00114-004-0592-y pdf

84. Byers, J.A. 2005b. Chemical constraints on the evolution of olfactory communication channels of moths. Journal of Theoretical Biology 235(2):199-206.
DOI: https://doi.org/10.1016/j.jtbi.2005.01.003 pdf

85. Byers, J.A. and Castle, S.J. 2005. Areawide models comparing synchronous versus asynchronous treatments for control of dispersing insect pests. Journal of Economic Entomology 98(6):1763-1773.
DOI: https://doi.org/10.1093/jee/98.6.1763 pdf

86. Byers, J.A. 2006a. Pheromone component patterns of moth evolution revealed by computer analysis of the Pherolist. Journal of Animal Ecology 75(2):399-407.
DOI: https://doi.org/10.1111/j.1365-2656.2006.01060.x pdf

87. Byers, J.A. 2006b. Analysis of insect and plant colors in digital images using Java software on the Internet. Annals of the Entomological Society of America 99(5):865-874.
DOI: https://doi.org/10.1603/0013-8746(2006)99%5B865:AOIAPC%5D2.0.CO;2 pdf

88. Byers, J.A. 2006c. Production and predator-induced release of volatile chemicals by the plant bug Lygus hesperus. Journal of Chemical Ecology 32(10):2205-2218.
DOI: https://doi.org/10.1007/s10886-006-9140-x pdf

89. El-Sayed, A.M., Suckling, D.M., Wearing, C.H., and Byers, J.A. 2006. Potential of mass trapping for long-term pest management and eradication of invasive species. Journal of Economic Entomology 99(5):1550-1564.
DOI: https://doi.org/10.1093/jee/99.5.1550 pdf

90. Thoss, V., and Byers, J.A. 2006. Monoterpene chemodiversity of ponderosa pine in relation to herbivory and bark beetle colonization. Chemoecology 16(1):51-58.
DOI: https://doi.org/10.1007/s00049-005-0331-7 pdf

91. Byers, J.A. 2007. Simulation of mating disruption and mass trapping with competitive attraction and camouflage. Environmental Entomology 36:1328-1338.
DOI: https://doi.org/10.1603/0046-225X(2007)36%5B1328:SOMDAM%5D2.0.CO;2 pdf

92. Natwick, E.T, Byers, J.A., Chu, C.C., Lopez, M., and Henneberry, T.J. 2007. Early detection and mass trapping of Frankliniella occidentalis and Thrips tabaci in vegetable crops. Southwestern Entomologist 32(4):229-238.
DOI: https://doi.org/10.3958/0147-1724-32.4.229 pdf

93. Blackmer, J.L., Byers, J.A., and Rodriguez-Saona, C. 2008. Evaluation of color traps for monitoring Lygus spp.: Design, placement, height, time of day, and non-target effects. Crop Protection 27(2):171-181.
DOI: https://doi.org/10.1016/j.cropro.2007.05.003 pdf

94. Byers, J.A. 2008. Active space of pheromone plume and its relationship to effective attraction radius in applied models. Journal of Chemical Ecology 34(9):1134-1145.
DOI: https://doi.org/10.1007/s10886-008-9509-0 pdf.
95. El-Sayed, A.M, Byers, J.A., Manning, L.M., Jürgens, A., Mitchell, V.J., and Suckling, D.M. 2008. Floral scent of Canada thistle and its potential as a generic insect attractant. Journal of Economic Entomology 101(3):720-727.
DOI: https://doi.org/10.1093/jee/101.3.720 pdf.

96. Byers, J.A. 2009. Modeling distributions of flying insects: Effective attraction radius of pheromone in two and three dimensions. Journal of Theoretical Biology 256(1):81-89.
DOI: https://doi.org/10.1016/j.jtbi.2008.09.002 pdf.

97. Blackmer, J.L., and Byers, J.A. 2009. Lygus spp. (Heteroptera: Miridae) host-plant interactions with Lesquerella fendleri (Brassicaceae), a new crop in the arid southwest. Environmental Entomology 38(1):159-167.
DOI: https://doi.org/10.1603/022.038.0120 pdf.

98. El-Sayed, A.M, Suckling, D.M, Byers, J.A., Jang, E.B., and Wearing, C.H. 2009. Potential of "lure and kill" in long-term pest management and eradication of invasive species. Journal of Economic Entomology 102(3):815-835.
DOI: https://doi.org/10.1603/029.102.0301 pdf.

99. Huang, C.H., Yan, F.M., Byers, J.A., Wang, R.J., and Xu, C.R. 2009. Volatiles induced by the larvae of the Asian corn borer (Ostrinia furnacalis) in maize plants affect behavior of conspecific larvae and female adults. Insect Science 16(4):311-320.
DOI: https://doi.org/10.1111/j.1744-7917.2009.01257.x pdf

100. Zada, A., Falach, L., and Byers, J.A. 2009. Development of sol-gel formulations for slow release of pheromones. Chemoecology 19(1):37-45.
DOI: https://doi.org/10.1007/s00049-009-0007-9 pdf.

101. Salvucci, M.E., Barta, C., Byers, J.A., and Canarini, A. 2010. Photosynthesis and assimilate partitioning between carbohydrates and isoprenoid products in vegetatively active and dormant guayule: physiological and environmental constraints on rubber accumulation in a semi-arid shrub. Physiologia Plantarum 140(4):368-379.
DOI: https://doi.org/10.1111/j.1399-3054.2010.01409.x pdf.

102. Brent, C.S. and Byers, J.A. 2011. Female attractiveness modulated by a male-derived antiaphrodisiac pheromone in a plant bug. Animal Behaviour 82(5):937-943.
DOI: https://doi.org/10.1016/j.anbehav.2011.08.010 pdf.

103. Byers, J.A. 2011. Analysis of vertical distributions and effective flight layers of insects: Three-dimensional simulation of flying insects and catch at trap heights. Environmental Entomology 40(5):1210-1222.
DOI: https://doi.org/10.1603/EN11043 pdf.

104. Byers, J.A. and Levi-Zada, A. 2011. Individual variation of (S)-4-methyl-3-heptanone in heads of braconid wasp, Leiophron uniformis, and Pogonomyrmex ants indicates costs of semiochemical production. Chemoecology 21(1):35-44.
DOI: https://doi.org/10.1007/s00049-010-0064-0 pdf.

105. Zhang, Q.H., Ma, J.H., Yang, Q.Q., Byers, J.A., Klein, M. G., Zhao, F.Y., and Luo, Y.Q. 2011. Olfactory and visual responses of the long-legged chafer Hoplia spectabilis Medvedev (Coleoptera: Scarabaeidae) in Qinghai province, China. Pest Management Science 67(2):162-169.
DOI: https://doi.org/10.1002/ps.2045 pdf.

106. Byers, J.A. 2012a. A population genetic model of evolution of host-mate attraction and nonhost repulsion in a bark beetle Pityogenes bidentatus. Psyche: A Journal of Entomology vol. 2012, ID 529573, pp. 1-9.
DOI: https://doi.org/10.1155/2012/529573 pdf.

107. Byers, J.A. 2012b. Estimating insect flight densities from attractive trap catches and flight height distributions. Journal of Chemical Ecology 38(5):592-601.
DOI: https://doi.org/10.1007/s10886-012-0116-8 pdf.

108. Byers, J.A. 2012c. Bark beetles, Pityogenes bidentatus, orienting to aggregation pheromone avoid conifer monoterpene odors when flying but not when walking. Psyche: A Journal of Entomology vol. 2012, ID 940962, pp. 1-10.
DOI: https://doi.org/10.1155/2012/940962 pdf.

109. Byers, J.A. 2012d. Modelling female mating success during mass trapping and natural competitive attraction of searching males or females. Entomologia Experimentalis et Applicata 145(3):228-237.
DOI: https://doi.org/10.1111/eea.12006 pdf
110. Byers, J.A. and G. Birgersson. 2012. Host-tree monoterpenes and biosynthesis of aggregation pheromones in the bark beetle Ips paraconfusus. Psyche: A Journal of Entomology vol. 2012, ID 539624, pp. 1-10.
DOI: https://doi.org/10.1155/2012/539624 pdf

111. Han, B., Zhang, Q.H., and Byers, J.A. 2012. Attraction of the tea aphid, Toxoptera aurantii, to combinations of volatiles and colors related to tea plants. Entomologia Experimentalis et Applicata 144(3):258-269.
DOI: https://doi.org/10.1111/j.1570-7458.2012.01303.x pdf

112. Lu, R.C., Wang, H.B., Zhang, Z., Byers, J.A., Jin, Y.J., Wen, H.F., Shi, W.J. 2012. Coexistence and competition between Tomicus yunnanensis and T. minor (Coleoptera: Scolytinae) in Yunnan pine. Psyche: A Journal of Entomology vol. 2012, ID 185312, pp. 1-6.
DOI: https://doi.org/10.1155/2012/185312 pdf

113. Lu, R.C., Wang, H.B., Zhang, Z., Byers, J.A., Jin, Y.J., Wen, H.F., and Shi, W.J. 2012. Attraction of Tomicus yunnanensis (Coleoptera: Scolytinae) to Yunnan pine logs with and without periderm or phloem: an effective monitoring bait. Psyche: A Journal of Entomology vol. 2012, ID 794683, pp. 1-5.
DOI: https://doi.org/10.1155/2012/794683 pdf

114. Rodriguez-Saona, C.R., Byers, J.A., and Schiffhauer, D. 2012. Effect of trap color and height on captures of blunt-nosed and sharp-nosed leafhoppers (Hemiptera: Cicadellidae) and non-target arthropods in cranberry bogs. Crop Protection 40:132-144.
DOI: https://doi.org/10.1016/j.cropro.2012.05.005 pdf

115. Byers, J.A., Birgersson, G., and Francke, W. 2013. Aggregation pheromones of bark beetles Pityogenes quadridens and P. bidentatus colonizing Scotch pine: olfactory avoidance of interspecific mating and competition. Chemoecology 23(4):251-261.
DOI: https://doi.org/10.1007/s00049-013-0139-9 pdf

116. Byers, J.A. 2013. Modeling and regression analysis of semiochemical dose-response curves of insect antennal reception and behavior. Journal of Chemical Ecology 39(8):1081-1089.
DOI: https://doi.org/10.1007/s10886-013-0328-6 pdf ... Supplementary Tables S1, S2 and S3 with References pdf

117. Byers, J.A., Fefer, D., and Levi-Zada, A. 2013. Sex pheromone component ratios and mating isolation among three Lygus plant bug species of North America. Naturwissenschaften 100(12):1115-1123.
DOI: https://doi.org/10.1007/s00114-013-1113-7 pdf

118. Byers, J.A. and Naranjo, S.E. 2014. Detection and monitoring of pink bollworm moths and invasive insects using pheromone traps and encounter rate models. Journal of Applied Ecology 51(4):1041-1049.
DOI: https://doi.org/10.1111/1365-2664.12270 pdf

119. Byers, J.A. 2014. Response to Martini and Habeck: Semiochemical dose-response curves fit by kinetic formation functions. Journal of Chemical Ecology 40(11):1165-1166.
DOI: https://doi.org/10.1007/s10886-014-0527-9 pdf

120. Byers, J.A. 2015. Earwigs (Labidura riparia) mimic rotting-flesh odor to deceive vertebrate predators. Science of Nature 102(article 38):1-10.
DOI: https://doi.org/10.1007/s00114-015-1288-1 pdf

Visiting Scientist / Affiliation, Department of Entomology, Hebrew University of Jerusalem, Rehovot, Israel:

121. El-Sayed, A.M., Byers, J.A., and Suckling, D.M. 2016. Pollinator-prey conflicts in carnivorous plants: When flower and trap properties mean life or death. Scientific Reports 6:21065. 11 pp.
DOI: https://doi.org/10.1038/srep21065 pdf ... Supplementary Material pdf

122. El-Sayed, A.M., Knight, A.L., Byers, J.A., Judd, G.J.R., and Suckling, D.M. 2016. Caterpillar-induced plant volatiles attract conspecific adults in nature. Scientific Reports 6:37555. 14 pp.
DOI: https://doi.org/10.1038/srep37555 pdf

123. Brent, C.S., Byers, J.A., and Levi-Zada, A. 2017. An insect anti-antiaphrodisiac. eLife 6:e24063. 15 pp.
DOI: https://doi.org/10.7554/eLife.24063 pdf

124. Byers, J.A., Maoz, Y., and Levi-Zada, A. 2017. Attraction of the Euwallacea sp. near fornicatus (Coleoptera: Curculionidae) to quercivorol and to infestations in avocado. Journal of Economic Entomology 110(4):1512-1517.
DOI: https://doi.org/10.1093/jee/tox151 pdf

125. Levi-Zada, A., Sadowsky, A., Dobrinin, S., Ticuchinski, T., David, M., Fefer, D., Dunkelblum, E., and Byers, J.A. 2018. Monitoring and mass-trapping methodologies using pheromones: The lesser date moth Batrachedra amydraula. Bulletin of Entomological Research 108(1):58-68.
DOI: https://doi.org/10.1017/S0007485317000487 pdf

126. Byers, J.A., Maoz, Y., Wakarchuk, D., Fefer, D., and Levi-Zada, A. 2018. Inhibitory effects of semiochemicals on the attraction of an ambrosia beetle Euwallacea nr. fornicatus to quercivorol. Journal of Chemical Ecology 44(6):565-575.
DOI: https://doi.org/10.1007/s10886-018-0959-8 pdf

127. Byers, J.A., Sadowsky, A, and Levi-Zada, A. 2018. Index of host habitat preference explored by movement-based simulations and trap captures. Journal of Animal Ecology 87(5):1320-1330.
DOI: https://doi.org/10.1111/1365-2656.12877 pdf

Research Consultant in Entomology - Chemical Ecology, Semiochemical Solutions, Beer Yaakov, Israel:

128. Levi–Zada, A., Levy, A., Rempoulakis, P., Fefer, D., Steiner, S., Gazit, Y., Nestel, D., Yuval, B., and Byers, J.A. 2020. Diel rhythm of volatile emissions of males and females of the peach fruit fly Bactrocera zonata. Journal of Insect Physiology 120:103970.
DOI: https://doi.org/10.1016/j.jinsphys.2019.103970 pdf

129. Byers, J.A., Maoz, Y., Fefer, D., and Levi–Zada, A. 2020. Semiochemicals affecting attraction of ambrosia beetle Euwallacea fornicatus (Coleoptera: Curculionidae: Scolytinae) to quercivorol: developing push-pull control. Journal of Economic Entomology 113(5):2120-2127.
DOI: https://doi.org/10.1093/jee/toaa127 pdf

130. Naranjo, S.E., Hagler, J.R., and Byers, J.A. 2021. Methyl salicylate fails to enhance arthropod predator abundance or predator to pest ratios in cotton. Environmental Entomology 50(2):293-305.
DOI: https://doi.org/10.1093/ee/nvaa175 pdf

131. Byers, J.A., Maoz, Y., Cohen, B., Golani, M., Fefer, D., and Levi-Zada, A. 2021. Protecting avocado trees from ambrosia beetles by repellents and mass trapping (push–pull): experiments and simulations. Journal of Pest Science 94(3):991-1002.
DOI: https://doi.org/10.1007/s10340-020-01310-x pdf

132. Levi-Zada, A., and Byers, J.A. 2021. Circadian rhythms of insect pheromone titer, calling, emission, and response: a review. Science of Nature 108(article 35):1-20.
DOI: https://doi.org/10.1007/s00114-021-01746-w pdf ... Supplementary Information: Levi-Zada and Byers 2021 Sci. Nat. Suppl. Info. Tables_Refs

133. Byers, J.A., and Levi-Zada, A. 2022. Modelling push-pull management of pest insects using repellents and attractive traps in fruit tree orchards. Pest Management Science 78(8):3630-3637.
DOI: https://doi.org/10.1002/ps.7005 pdf ... You may download Java JAR simulation executible: pushpull2.jar and two image files: pshb1.png and pushpull2a50.jpg note: pushpull2.jar will download as tab at lower left, click on [ ˄ ] and "Show in folder" to save in a directory and run with double click from Windows Explorer, or click "Open" to run directly

134. Ahmed, D.A., Beidas, A., Petrovskii, S.V., Bailey, J.D., Bonsall, M.B., Hood, A.S.C., Byers, J.A., Hudgins, E.J., Russell, J.C., Ružicková, J., Bodey, T.W., Renault, D., Bonnaud, E., Haubrock, P.J., Soto, I., and Haase, P. 2023. Capture efficiency of pitfall traps based on sampling strategy and the movement of ground-dwelling arthropods. Methods in Ecology and Evolution 14(11):2827-2843.
DOI: https://doi.org/10.1111/2041-210X.14174 pdf

135. Levi-Zada, A., Steiner, S., Fefer, D., and Byers, J.A. 2025. Repelling fruit flies with essential oils and their components: the peach fruit fly Bactrocera zonata. Journal of Chemical Ecology 51:81.
DOI: https://doi.org/10.1007/s10886-025-01628-9 pdf ... Supplementary Information: Levi-Zada et al. ESM J. Chem. Ecol. 2025.pdf

Other Publications (Book Chapters, Journal Editorials, Israeli Journals, and Internet Webpages):

1. Schlyter, F., Byers, J.A., Löfqvist, J., Leufvén, A., and Birgersson, G. 1988. Reduction of attack density of the bark beetles Ips typographus and Tomicus piniperda on host bark by verbenone inhibition of attraction to pheromone and host kairomone. pp. 53-68. In: Payne, T.L. and Saarenmaa, H. (eds.), Integrated Control of Scolytid Bark Beetles, Virginia Tech. Press, Blacksburg. 355 pp. pdf

2. Byers, J.A. 1995. Host tree chemistry affecting colonization in bark beetles, in R.T. Cardé and W.J. Bell (eds.). Chemical Ecology of Insects 2. Chapman and Hall, New York, pp. 154-213. DOI: https://doi.org/10.1007/978-1-4615-1765-8_5 pdf

3. Byers, J.A. 2004b. Chemical ecology of bark beetles in a complex olfactory landscape. pp. 89-134. In, Bark and Wood Boring Insects in Living Trees in Europe, a Synthesis, F. Lieutier, K.R. Day, A. Battisti, J.C. Grégoire, H. Evans (Eds). Kluwer Academic Publishers, Dordrecht, The Netherlands. pdf

4. Byers, J.A. and Zhang, Q.H. 2012. Chemical ecology of bark beetles in regard to search and selection of host trees, pp. 150-190. In, Recent Advances in Entomological Research. (eds.) T.-X. Liu and L. Kang. The Higher Education Press, Beijing, and Springer-Verlag GmbH, Berlin. pdf

5. Byers, J.A. 2012. Ecological interactions of bark beetles with host trees. Editorial Article. Psyche: A Journal of Entomology Article ID 252961, 3 pages. pdf

6. Levi-Zada, A., David, M., Fefer, D., Dunkelblum, E., Sadowsky, A., Ticuchinski, T., Katzman, A., Yossel, E., Dobrinin, S., and Byers, J.A. 2019. Lesser date moth: Development of mass trapping tools by pheromones. (in Hebrew) Alon Hanotea 73(2):20-25. pdf

7. Byers, J.A., Maoz, Y., Fefer, D., Levi-Zada, A. 2020. Protecting avocado trees from ambrosia beetles by push-pull method. (in Hebrew) Alon Hanotea 74(1):38-42. pdf

Popular Science and Education:

My web site previewed as "Bug juice" in
Science 288:767.
on 5 May 2000.
 
Theses:

Byers, J.A. 1973. Plant growth hormones in pinyon insect galls. Master of Science, Department of Entomology, Colorado State University,
     Fort Collins, Colorado, U.S.A.  pdf

Byers, J.A. 1978. Mutual inhibition of the attractive response to pheromones of Dendroctonus brevicomis and Ips paraconfusus,
     and the physiology of pheromone production in Ips paraconfusus. Doctor of Philosophy, Department of Entomological Sciences,
	 University of California, Berkeley, California, U.S.A.  pdf

Miscellaneous Letters in Entomological Publications:

Byers, J.A. 1983. (Letter proposing new journal called `Entomology'). ESA Newsletter (Entomological Society of America) 6(10):1.
      pdf

Byers, J.A. 1984. (Letter proposing name change of `Ann. Entomol. Soc. Amer.' to `Entomology', also proposes free page charges to increase
     competition and quality in ESA journals). ESA Newsletter 7(8):6.  pdf

Byers, J.A. 1985. (Letter discussing International membership, a recent questionnaire, `Entomology' journal, and the need for having
     national meetings in different regions). ESA Newsletter 8(10):3.  pdf

Byers, J.A. 1987. What's in a name? Bulletin of the Entomological Society of America 33:208.
and (c) by John A. Byers ¥